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The base of the mapped sections consists of a well developed, thick, aeolian facies (8). U.) is composed of poorly cemented sand and argillaceous sandstone alternation characterized by dense networks of root tubules/root molds (palaeosols) and termite nests (9, 10). The uniform stratigraphy at the TM localities allowed us to use absolute ages from both TM 266, where Toumaï was discovered, and TM 254 to assign an age to Toumaï.

It was hoped that the tephra layer would contain material datable by the Be associates with continental particles, where it decays with a half-life of ≈1.4 million years.

Five decades have passed since the proposal of the molecular clock hypothesis, which states that the rate of evolution at the molecular level is constant through time and among species.

This hypothesis has become a powerful tool in evolutionary biology, making it possible to use molecular sequences to estimate the geological ages of species divergence events.

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With recent advances in Bayesian clock dating methodology and the explosive accumulation of genetic sequence data, molecular clock dating has found widespread applications, from tracking virus pandemics and studying the macroevolutionary process of speciation and extinction to estimating a timescale for life on Earth.

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